［1］YANG H,MATSUBAYASHI Y,NAKAMURA K,et al.Diversity of Arabidopsis genes encoding precursors for phytosulfokine,a peptide growth factor［J］.Plant Physiology,2001,127(3):842-851.［2］SAUTER M.Phytosulfokine peptide signalling［J］.Journal of Experimental Botany,2015,66(17):5161-5169.［3］MATSUBAYASHI Y,SAKAGAMI Y.Peptide hormones in plants［J］.Annual Review of Plant Biology,2006,57:649-674.［4］LIU Y,ZHANG D,LI M,et al.Overexpression of PSK-γ in Arabidopsis promotes growth without influencing pattern-triggered immunity［J］.Plant Signaling & Behavior,2019,14(12):1684423.［5］MATSUBAYASHI Y,SAKAGAMI Y.Phytosulfokine,sulfated peptides that induce the proliferation of single mesophyll cells of Asparagus officinalis L［J］.Proceedings of the National Academy of Sciences of the United States of America,1996,93(15):7623-7627.［6］YU L,DI Q,ZHANG D,et al.A legume-specific novel type of phytosulfokine,PSK-δ,promotes nodulation by enhancing nodule organogenesis［J］.Journal of Experimental Botany,2022,73(8):2698-2713.［7］SRIVASTAVA R,LIU J X,HOWELL S H.Proteolytic processing of a precursor protein for a growth-promoting peptide by a subtilisin serine protease in Arabidopsis［J］.The Plant Journal:for Cell and Molecular Biology,2008,56(2):219-227.［8］KOMORI R,AMANO Y,OGAWA-OHNISHI M,et al.Identification of tyrosylprotein sulfotransferase in Arabidopsis［J］.Proceedings of the National Academy of Sciences of the United States of America,2009,106(35):15067-15072.［9］YANG H,MATSUBAYASHI Y,NAKAMURA K,et al.Oryza sativa PSK gene encodes a precursor of phytosulfokine-alpha,a sulfated peptide growth factor found in plants［J］.Proceedings of the National Academy of Sciences of the United States of America,1999,96(23):13560-13565.［10］MATSUBAYASHI Y,TAKAGI L,SAKAGAMI Y.Phytosulfokine-alpha,a sulfated pentapeptide,stimulates the proliferation of rice cells by means of specific high-and low-affinity binding sites［J］.Proceedings of the National Academy of Sciences of the United States of America,1997,94(24):13357-13362.［11］CHEN J Y,YANG Y H,XU H X,et al.Stimulatory effect of phytosulfokine-alpha on the proliferation of tobacco ‘Bright Yellow 2’ suspension cells［J］.Zhi Wu Sheng Li Yu Fen Zi Sheng Wu Xue Xue Bao=Journal of Plant Physiology and Molecular Biology,2007,33(1):39-45.［12］KIEKOWSKA A,ADAMUS A.Peptide growth factor phytosulfokine-α stimulates cell divisions and enhances regeneration from B.oleracea var.capitata L.protoplast culture［J］.Journal of Plant Growth Regulation,2019,38(3):931-944.［13］MATSUBAYASHI Y,GOTO T,SAKAGAMI Y.Chemical nursing:Phytosulfokine improves genetic transformation efficiency by promoting the proliferation of surviving cells on selective media［J］.Plant Cell Reports,2004,23(3):155-158.［14］YAMAKAWA S,MATSUBAYASHI Y,SAKAGAMI Y,et al.Promotive effects of the peptidyl plant growth factor,phytosulfokine-alpha,on the growth and chlorophyll content of Arabidopsis seedlings under high night-time temperature conditions［J］.Bioscience,Biotechnology,and Biochemistry,1999,63(12):2240-2243.［15］SONG H,WANG X,HU W,et al.A cold-induced phytosulfokine peptide is related to the improvement of loquat fruit chilling tolerance［J］.Food Chemistry,2017,232:434-442.［16］LOIVAMEKI M,STHRWOHLDT N,DEEKEN R,et al.A role for PSK signaling in wounding and microbial interactions in Arabidopsis［J］.Physiologia Plantarum,2010,139(4):348-357.［17］IGARASHI D,TSUDA K,KATAGIRI F.The peptide growth factor,phytosulfokine,attenuates pattern-triggered immunity［J］.The Plant Journal:for Cell and Molecular Biology,2012,71(2):194-204.［18］MOSHER S,SEYBOLD H,RODRIGUEZ P,et al.The tyrosine-sulfated peptide receptors PSKR1 and PSY1R modify the immunity of Arabidopsis to biotrophic and necrotrophic pathogens in an antagonistic manner［J］.The Plant Journal:for Cell and Molecular Biology,2013,73(3):469-482.［19］CHEN Y F,MATSUBAYASHI Y,SAKAGAMI Y.Peptide growth factor phytosulfokine-α contributes to the pollen population effect［J］.Planta,2000,211(5):752-755.［20］STHRWOHLDT N,DAHLKE R I,KUTSCHMAR A,et al.Phytosulfokine peptide signaling controls pollen tube growth and funicular pollen tube guidance in Arabidopsis thaliana［J］.Physiologia Plantarum,2015,153(4):643-653.［21］KOU X,LIU Q,SUN Y,et al.The peptide PbrPSK2 from phytosulfokine family induces reactive oxygen species (ROS) production to regulate pear pollen tube growth［J］.Frontiers in Plant Science,2020,11:601993.［22］YU L,LIU Y,LIU Y,et al.Overexpression of phytosulfokine-α induces male sterility and cell growth by regulating cell wall development in Arabidopsis［J］.Plant Cell Reports,2016,35(12):2503-2512.［23］WANG C,YU H,ZHANG Z,et al.Phytosulfokine is involved in positive regulation of Lotus japonicus nodulation［J］.Molecular Plant-Microbe Interactions:MPMI,2015,28(8):847-855.［24］AGHDAM M S,ALIKHANI-KOUPAEI M.Exogenous phytosulfokine α (PSKα) applying delays senescence and relief decay in strawberry fruits during cold storage by sufficient intracellular ATP and NADPH availability［J］.Food Chemistry,2021,336:127685.［25］AGHDAM M S,SAYYARI M,LUO Z.Exogenous application of phytosulfokine α (PSKα) delays yellowing and preserves nutritional quality of broccoli florets during cold storage［J］.Food Chemistry,2020,333:127481.［26］AGHDAM M S,EBRAHIMI A,SHEIKH-ASSADI M,et al.Endogenous phytosulfokine α (PSKα) signaling delays petals senescence and prolongs vase life of cut rose flowers (Rosa hybrida cv.Angelina)［J］.Scientia Horticulturae,2021,289:110444.［27］AMANO Y,TSUBOUCHI H,SHINOHARA H,et al.Tyrosine-sulfated glycopeptide involved in cellular proliferation and expansion in Arabidopsis［J］.Proceedings of the National Academy of Sciences of the United States of America,2007,104(46):18333-18338.［28］MATSUBAYASHI Y,OGAWA M,MORITA A,et al.An LRR receptor kinase involved in perception of a peptide plant hormone,phytosulfokine［J］.Science,2002,296(5572):1470-1472.［29］ZHANG H,HU Z,LEI C,et al.A plant phytosulfokine peptide initiates auxin-dependent immunity through cytosolic Ca2+ signaling in tomato［J］.The Plant Cell,2018,30(3):652-667.［30］KWEZI L,RUZVIDZO O,WHEELER J I,et al.The phytosulfokine (PSK) receptor is capable of guanylate cyclase activity and enabling cyclic GMP-dependent signaling in plants［J］.The Journal of Biological Chemistry,2011,286(25):22580-22588.［31］KWEZI L,MEIER S,MUNGUR L,et al.The Arabidopsis thaliana brassinosteroid receptor (AtBRI1) contains a domain that functions as a guanylyl cyclase in vitro［J］.PLoS One,2007,2(5):e449.［32］QI Z,VERMA R,GEHRING C,et al.Ca2+ signaling by plant Arabidopsis thaliana Pep peptides depends on AtPepR1,a receptor with guanylyl cyclase activity,and cGMP-activated Ca2+ channels［J］.Proceedings of the National Academy of Sciences of the United States of America,2010,107(49):21193-21198.［33］LADWIG F,DAHLKE R I,STHRWOHLDT N,et al.Phytosulfokine regulates growth in Arabidopsis through a response module at the plasma membrane that includes CYCLIC NUCLEOTIDE-GATED CHANNEL17,H+-ATPase,and BAK1［J］.The Plant Cell,2015,27(6):1718-1729.［34］MA Y,WALKER R K,ZHAO Y,et al.Linking ligand perception by PEPR pattern recognition receptors to cytosolic Ca2+ elevation and downstream immune signaling in plants［J］.Proceedings of the National Academy of Sciences of the United States of America,2012,109(48):19852-19857.［35］ZHAO Y,QI Z,BERKOWITZ G A.Teaching an old hormone new tricks:Cytosolic Ca2+ elevation involvement in plant brassinosteroid signal transduction cascades［J］.Plant Physiology,2013,163(2):555-565.［36］MATSUBAYASHI Y,MORITA A,MATSUNAGA E,et al.Physiological relationships between auxin,cytokinin,and a peptide growth factor,phytosulfokine-α,in stimulation of asparagus cell proliferation［J］.Planta,1999,207(4):559-565.［37］EUN C H,KO S M,MATSUBAYASHI Y,et al.Phytosulfokine-α requires auxin to stimulate carrot non-embryogenic cell proliferation［J］.Plant Physiology and Biochemistry,2003,41(5):447-452.［38］ZHOU W,WEI L,XU J,et al.Arabidopsis tyrosylprotein sulfotransferase acts in the auxin/PLETHORA pathway in regulating postembryonic maintenance of the root stem cell niche［J］.The Plant Cell,2010,22(11):3692-3709.［39］HARTMANN J,STHRWOHLDT N,DAHLKE R I,et al.Phytosulfokine control of growth occurs in the epidermis,is likely to be non-cell autonomous and is dependent on brassinosteroids［J］.The Plant Journal:for Cell and Molecular Biology,2013,73(4):579-590.［40］WU T,KAMIYA T,YUMOTO H,et al.An Arabidopsis thaliana copper-sensitive mutant suggests a role of phytosulfokine in ethylene production［J］.Journal of Experimental Botany,2015,66(13):3657-3667.［41］杨艳会.新型肽类生长调节物质:植物磺肽素(Phytosulfokine)的研究［D］.郑州:河南农业大学,2006.［42］王雅男,徐晨,李桂新,等.植物多肽类调节物质磺肽素(PSK)的功能与作用机制［J］.植物生理学报,2022,58(10):1818-1828.［43］师恺,方瀚墨,马巧梅,等.植物磺肽素在提高农作物果实风味和营养品质中的应用［P］.中国:CN114651622A，2022-06-24.［44］喻景权,师恺,张欢,等.植物磺肽素-α在提高植物灰霉病抗性中的应用［P］.中国:CN107114369B，2020-01-21.［45］师恺,张欢,史军营,等.植物磺肽素-α在提高植物黄化曲叶病毒病抗性中的应用［P］.中国:CN105900983B，2018-03-16.