[1]PINHEIRO T T,NISHIMURA D S,DE NADAI F B,et al.Selection of reference genes for expression analyses of red-fleshed sweet orange (Citrus sinensis)[J].Genetics and Molecular Research,2015,14(4):18440-18451.[2]RUMAINUM I M,WORARAD K,YAMAKI Y,et al.Effects of developmental stages,light,and an auxin polar transport inhibitor on the skin and flesh pigmentation of red-fleshed peach fruit[J].Horticulture,2016,85(2):141-147.[3]SEKIDO K,HAYASHI Y,YAMADA K,et al.Efficient breeding system for red-fleshed apple based on linkage with S3-RNase allele in ‘Pink Pearl’[J].HortScience,2010,45(4):534-537.[4]SADILOVA E,STINTZING F C,CARLE R.Chemical quality parameters and anthocyanin pattern of red-fleshed Weirouge apples[J].Journal of Applied Botany and Food Quality,2006,80(1):82-87.[5]BALAZS A,TTH M,BLAZICS B,et al.Investigation of dietary important components in selected red fleshed apples by GC-MS and LC-MS[J].Fitoterapia,2012,83(8):1356-1363.[6]WANG N,JIANG S H,ZHANG Z Y,et al.Malus sieversii:The origin,flavonoid synthesis mechanism,and breeding of red-skinned and red-fleshed apples[J].Horticulture Research,2018(5):70.[7]RUPASINGHE H P V,HUBER G M,EMBREE C,et al.Red-fleshed apple as a source for functional beverages[J].Canadian Journal of Plant Science,2010,90(1):95-100.[8]JEFFERSON R M.History,progeny,and locations of crabapple of documented authentic origin[M].Washington D C:Agricultural Research Service,U.S.Dept of Agriculture,1965.[9]陆秋农,贾定贤.中国果树志.苹果卷[M].北京:中国农业科技出版社,中国林业出版社,1999.[10]PENG T,MORIGUCHI T.The molecular network regulating the coloration in apple[J].Scientia Horticulturae,2013,163:1-9.[11]李育农.苹果属植物种质资源研究[M].北京:农业出版社,2001.[12]van NOCKER S V,BERRY G,NAJDOWSKI J,et al.Genetic diversity of red-fleshed apples (Malus)[J].Euphytica,2011,185:281-293.[13]WRDIG J,FLACHOWSKY H,HFER M,et al.Phenotypic and genetic analysis of the German Malus Germplasm Collection in terms of type 1 and type 2 red-fleshed apples[J].Gene,2014,544(2):198-207.[14]UMEMURA H,SHIRATAKE K,MATSUMOTO S,et al.Practical breeding of Red-fleshed Apple:Cultivar combination for efficient red-fleshed progeny production[J].HortScience,2011,46(8):1098-1101.[15]VOLZ R K,ORAGUZIE N C,WHITWORTH C J,et al.Breeding for red flesh colour in apple:Progress and challenges[J].Acta Horticulturae,2009,814:337-342.[16]VOLZ R K,KUMAR S,CHAGN D,et al.Genetic relationship between red flesh and fruit quality traits in apple[J].Acta Horticulturae,2013,976:363-368.[17]ESPLEY R V,HELLENS R P,PUTTERILL J,et al.Red colouration in apple fruit is due to the activity of the MYB transcription factor,[STBX]MdMYB10[STBZ][J].Plant Journal,2007,49(3):414-427.[18]BARS-CORTINA D,MACI A,IGLESIAS I,et al.Phytochemical profiles of new red-fleshed apple varieties compared with traditional and new white-fleshed varieties[J].Journal of Agricultural and Food Chemistry,2017,65(8):1684-1696.[19]MALEC M,LE QUERE J M,SOTIN H,et al.Polyphenol profiling of a red-fleshed apple cultivar and evaluation of the color extractability and stability in the juice[J].Journal of Agricultural and Food Chemistry,2014,62(29):6944-6954.[20]陈学森,张晶,刘大亮,等.新疆红肉苹果杂种一代的遗传变异及功能型苹果优株评价[J].中国农业科学,2014,47(11):2193-2204.[21]许海峰,王楠,姜生辉,等.新疆红肉苹果杂种一代4个株系类黄酮含量及其合成相关基因表达分析[J].中国农业科学,2016,49(16):3174-3187.[22]MAZZA G,VELIOGLU Y S.Anthocyanins and other phenolic-compounds in fruits of red-flesh apples[J].Food Chemistry,1992,43(2):113-117.[23]SADILOVA E,STINTZING F C,CARLE R.Chemical quality parameters and anthocyanin pattern of red-fleshed Weirouge apples[J].Journal of Applied Botany and Food Quality,2006,80:82-87.[24]张翔,孙晓红,柏素花,等.4种红肉苹果提取液花青苷含量及体外抗氧化研究[J].青岛农业大学学报(自然科学版),2018,35(3):179-186.[25]路滨键,于克可,张玉刚,等.红肉苹果营养成分及生物活性物质分析[J].青岛农业大学学报(自然科学版),2018,35(1):9-15.[26]SATO H,OTAGAKI S,SAELAI P,et al.Varietal differences in phenolic compounds metabolism of type 2 red-fleshed apples[J].Scientia Horticulturae,2017,219:1-9.[27]FARAMARZI S,PACIFICO S,YADOLLAHI A,et al.Red-fleshed apples:Old autochthonous fruits as a novel source of anthocyanin antioxidants[J].Plant Foods for Human Nutrition,2015,70(3):324-330.[28]TOUFEKTSIAN M C,LORGERIL M D,NAGY N,et al.Chronic dietary intake of plant-derived anthocyanins protects the rat heart against is chemia-reperfusion injury[J].The Journal of Nutrition,2008,138:747-752.[29]安建平,宋来庆,赵玲玲,等.苹果愈伤组织超表达[STBX]MdNAC029[STBZ]促进花青苷积累[J].园艺学报,2018,45(5):845-854.[30]TAKOS A M,JAFF F W,JACOB S R,et al.Light-induced expression of a MYB gene regulates anthocyanin biosynthesis in red apples[J].Plant Physiology,2006,142(3):1216-1232.[31]BAN Y,HONDA C,HATSUYAMA Y,et al.Isolation and functional analysis of a MYB transcription factor gene that is a key regulator for the development of red coloration in apple skin[J].Plant and Cell Physiology,2007,48(7):958-970.[32]MORIMOTO T,HIRAMATSU Y,BANNO K.A major QTL controlling earliness of fruit maturity linked to the Red leaf/Red flesh trait in apple cv.‘Maypole’[J].Journal of the Japanese Society for Horticultural Science,2013,82(2):97-105.[33]UMEMURA H,OTAGAKI S,WADA M,et al.Expression and functional analysis of a novel MYB gene,[STBX]MdMYB110a_JP[STBZ],responsible for red flesh,not skin color in apple fruit[J].Planta,2013,238:65-76.[34]CHAGN D,LIN-WANG K,ESPLEY R V,et al.An ancient duplication of apple MYB transcription factors is responsible for novel red fruit-flesh phenotypes[J].Plant Physiology,2013,161(1):225-239.[35]ESPLEY R V,BOVY A,BAVA C,et al.Analysis of genetically modified red-fleshed apples reveals effects on growth and consumer attributes[J].Plant Biotechnology,2013,11(4):408-419.[36]TELIAS A,LIN-WANG K,STEVENSON D E,et al.Apple skin patterning is associated with differential expression of [STBX]MYB10[STBZ][J].BMC Plant Biology,2011,11(1):93.[37]XIE X B,LI S,ZHANG R F,et al.The bHLH transcription factor [STBX]MdbHLH3[STBZ] promotes anthocyanin accumulation and fruit colouration in response to low temperature in apples[J].Plant,Cell and Environment,2012,35(11):1884-1897.[38]PENG T,SAITO T,HONDA C,et al.Screening of UV-B-induced genes from apple peels by SSH:Possible involvement of MdCOP1-mediated signaling cascade genes in anthocyanin accumulation[J].Physiologia Plantarum,2013,148(3):432-444.[39]VIMOLMANGKANG S,ZHENG D M,HAN Y P,et al.Transcriptome analysis of the exocarp of apple fruit identifies light-induced genes involved in red color pigmentation[J].Gene,2014,534(1):78-87.[40]WANG N,ZHANG Z Y,JIANG S H,et al.Synergistic effects of light and temperature on anthocyanin biosynthesis in callus cultures of red-fleshed apple (Malus sieversii f.niedzwetzkyana)[J].Plant Cell,Tissue and Organ Culture,2016,127(1):217-227.[41]HONDA C,IWANAMI H,NARAMOTO K,et al.Thinning and bagging treatments and the growing region influence anthocyanin accumulation in red-fleshed apple fruit[J].Horticulture,2017,86(3):291-299.[42]MATSUMOTO K,KOBAYASHI T,KOUGO T,et al.Sunlight differentially affects the fruit skin,flesh,and core coloration of the type 2 red-fleshed apple ‘Kurenainoyume’:Optimization of fruit bagging treatment[J].Horticulture,2018,87(4):462-473.[43]ZHANG Y G,ZHAO R X,LIU W L,et al.The anthocyanins component and the influence factors of contents in red flesh apple ‘Hong-Xun No.1’[J].European Journal of Horticultural Science,2016,81(5):248-254.[44]HAMADA Y,SATO H,OTAGAKI S,et al.Breeding depression of red flesh apple progeny containing both functional [STBX]MdMYB10 and MYB110a_JP[STBZ] genes[J].Plant Breeding,2015,134(2):239-246.[45]YOSHIIE K.Apple tree named ‘Moon Rouge’[P].US,0219767,2016.[46]ZHANG X,HUO H,SUN X,et al.Nanocrystallization of anthocyanin extract from red-fleshed apple ‘QN-5’ improved its antioxidant effect through enhanced stability and activity under stressful conditions[J].Molecules 2019,24(7):180-182.[47]KOBELT M.Apple tree named‘Lurechild’[P].US,0150695,2016.[48]PARVANEH T,ABEDI B,DAVARYNEJAD G H,et al.Enzyme activity,phenolic and flavonoid compounds in leaves of Iranian red flesh apple cultivars grown on different rootstocks[J].Scientia Horticulturae,2019,246:862-870.