[1]NOLD R.Columbines:Aquilegia,Paraquilegia and semi-Aquilegia[M].Portland:Timber Press,2003. [2]FU D Z,ROBINSON O R.Aquilegia[M]//WU Z Y,RAVEN P (ed).Flora of China.Beijing:Science Press,2001. [3]WHITTALL J B,HODGES S A.Pollinator shifts drive increasingly long nectar spurs in columbine flowers[J].Nature,2007,447:706-710. [4]TANG L L,YU Q,SUN J F,et al.Floral traits and isolation of three sympatric Aquilegia species in the Qinling Mountains,China[J].Plant Syst Evol,2007,267:121-128. [5]BASTIDA J M,ALCNTARA J M,REY P J,et al.Extended phylogeny of Aquilegia:the biogeographical and ecological patterns of two simultaneousbut contrasting radiations[J].Plant Syst Evol,2010,284:171-185. [6]KRAMER E M.Aquilegia:A new model for plant development,ecology,and evolution[J].Annu Rev Plant Biol,2009,60:261-277. [7]KRAMER E M,HODGES S A.Aquilegiaas a model system for the evolution and ecology of petals[J].Philos Trans Roy Soc Lond Ser B,2010,365:477-490. [8]YEH F C,YANG R C,BOYLE T,et al.The user-friendly shareware for population genetic analysis[M].Canada:University of Alberta Press,1997. [9]EXCOFFIER L,LAVAL G,SCHNEIDER S.Arlequin ver.3.0:An integrated software package for population genetics data analysis[J].Evol Bioinform,2005(1):47-50. [10]ITAGAKI T,KIMURA M K,MAKI M.Differential self-fertilization rates in response to variation in floral traits within inflorescences of Aquilegia buergeriana var.oxysepala (Ranunculaceae)[J].Botanical Journal of the Linnean Society,2016,181(2):294-304. [11]〖JP〗MEDRANO M,CASTELLANOS M C,HERRERA C M.Comparative floraland vegetative differentiation between two European Aquilegiataxa along a narrow contact zone[J].Plant Syst Evol,2007,262:209-224. [12]GAFTA D,MUNCACIU S,CSERGO A M.Morphometric variation in a rare endemic Aquilegia(Ranunculaceae) in the Carpathians[J].Plant Biosyst,2006,140:297-306. [13]LAVERGNE S,DEBUSSCHE M,THOMPSON J D.Limitations onreproductive success in endemic Aquilegia viscosa(Ranunculaceae) relative to its widespread congener Aquilegia vulgaris:the interplay of herbivory and pollination[J].Oecologia,2005,142:212-220. [14]ITAGAKI T,KIMURA M K,LIAN C,et al.Development of microsatellite markers for Aquilegia buergeriana var.oxysepala (Ranunculaceae),a vulnerable Japanese herb[J].Plant Species Biology,2015,30:159-162. [15]ERST A S,VAULIN O V.Phylogenetic relationships among North Asian species of the genus Aquilegia based on molecular markers[J].Russian Journal of Genetics(Applied Research),2014,4(1):35-42. [16]FIOR S,LI M,OXELMAN B,et al.Spatiotemporal reconstruction of the Aquilegia rapid radiation through next generation sequencing of rapidly evolving cpDNA regions[J].New Phytol,2013,198(2):579-592. [17]SHAW J,LICKEY E B,SCHILLING E E,et al.Comparison of whole chloroplast genome sequences tochoose noncoding regions for phylogenetic studies inangiosperms:the tortoise and the hare III[J].Am J Bot,2007,94(3):275-288. [18]朱蕊蕊,高亦珂,张启翔.部分耧斗菜种的 AFLP 遗传多样性分析[C].First Inernational Symposium on Biomedicine and Engineering,2011:179-182. [19]朱蕊蕊,高亦珂,张启翔.耧斗菜属AFLP体系的建立与优化[J].华北农学报,2010,25(suppl):38-41. [20]朱蕊蕊,高亦珂,张启翔.耧斗菜属的物种进化与遗传学[J].分子植物育种,2010,8(4):784-789.