[1]JOHNSON J D.Do carotenoids serve as transmembrane radical channels?[J].Free Radical Biology and Medicine,2009,47:321-323. [2]POLIVKA T,FRANK H A.Molecular factors controlling photosynthetic light harvesting by carotenoids[J].Accounts of Chemical Research,2010,43:1125-1134. [3]CAZZONELLI C I,POGSON B J.Source to sink:Regulation of carotenoid biosynthesis in plants[J].Trends Plant Science,2010(15):266-274. [4]XIE X,YONEYAMA K.The strigolactone story[J].Annual Review of Phytopathology,2010,48:93-117. [5]OHMIYA A.Qualitative and quantitative control of carotenoid accumulation in flower petals[J].Scientia Horticulturae,2013,163:10-19. [6]EISENREICH W,BACHER A,ARIGONI D,et al.Biosynthesis of isoprenoids via the non-mevalonate pathway[J].Cellular and Molecular Life Sciences,2004,61:1401-1426. [7]KIM M J,KIM J K,KIM H J,et al.Genetic modification of the soybean to enhance the β-carotene content through seed-specific expression[J].PLoS One,2012,7(10):e48287. [8]ISAACASON T,OHAD I,BEYER P,et al.Analysis in vitro of the enzyme CRTISO establishes a poly-cis-carotenoid biosynthesis pathway in plants[J].Plant Physiology,2004,136:4246-4255. [9]CUNNINGHAM F X,POGSON B,SUN Z R,et al.Functional analysis of the beta and epsilon lycopene cyclase enzymes of Arabidopsis reveals a mechanism for control of cyclic carotenoid formation[J].Plant Cell,1996(8):1613-1626. [10]RONEN G,COHEN M,ZAMIR D,et al.Regulation of carotenoid biosynthesis during tomato fruit development:expression of the gene for lycopene epsilon-cyclase is downregulated during ripening and is elevated in the mutant delta[J].Plant Journal,1999(17):341-351. [11]CUNNINGHAM F X,GANTT E.One ring or two? Determination of ring number in carotenoids by lycopene epsilon-cyclases[J].Proceedings of the National Academy of Sciences,2001,98(5):2905-2910. [12]FORSTER B,POGSON B J,OSMOND C B.Lutein from deepoxidation of lutein epoxide replaces zeaxanthin to sustain an enhanced capacity for nonphotochemical chlorophyll fluorescence quenching in avocado shade leaves in the dark[J].Plant Physiology,2011,156:393-403. [13]ZHU C,YAMAMURA S,NISHIHARA M,et al.cDNAs for the synthesis of cyclic carotenoids in petals of Gentiana lutea and their regulation during flower development[J].Biochimica et Biophysica Acta,2003,1625:305-309. [14]JEKNIC Z,MORRE J T,JEKNIC S,et al.Cloning and functional characterization of a gene for capsanthin-capsorubin synthase from tiger lily(Lilium lancifolium Thunb.‘ Splendens ’)[J].Biochimica et Biophysica Acta,2012,53:1899-1912. [15]GUZMAN I,HAMBY S,ROMERO J.et al.Variability of carotenoid biosynthesis in orange colored Capsicum spp.[J].Plant Sci,2010,179:49-59. [16]WALTER M H,STRACK D.Carotenoids and their cleavage products:biosynthesis and functions[J].Natural Product Reports,2011(28):663-692. [17]TAN B C,JOSEPH L M,DENG W T,et al.Molecular characterization of the Arabidopsis 9-cis epoxycarotenoid dioxygenase gene family[J].Plant Journal,2003,35:44-56. [18]SCHMIDT H,KURTZER R,EISENREICH W,et al.Molecular characterization of the Arabidopsis 9-cis epoxycarotenoid dioxygenase gene family[J].Journal of Biological Chemistry,2006,281(15):9845-9851. [19]SIMKIN A J,SCHWARTZ S H,AULDRIDGE M,et al.The tomato carotenoid cleavage dioxygenase 1 genes contribute to the formation of the flavor volatiles β-ionone,pseudoionone,and geranylacetone[J].Plant Journal,2004,40:882-892. [20]SIMKIN A J,UNDERWOOD B A,AULDRIDGE M,et al.Circadian regulation of the PhCCD1 carotenoidcleavage dioxygenase controls emission of β-ionone,a fragrance volatile of petunia flowers[J].Plant Physiology,2004,136:3504-3514. [21]ADAMI M,DE F P,BRANDI F,et al.Identifying a carotenoid cleavage dioxygenase(ccd4)gene controlling yellow/white fruit flesh color of peach[J].Plant Molecular Biology Reporter,2013,31(5):1166-1175. [22]OHMIYA A,KISHIMOTO S,AIDA R,et al.Carotenoid cleavage dioxygenase(CmCCD4a)contributes to white color formation in chrysanthemum petals[J].Plant Physiology,2006,142:1193-1201. [23]RUYTER-SPIRA C,AL-BABILI S,VAN DER KROL S,et al.The biology of strigolactones[J].Trends in Plant Science,2013(18):72-83. [24]SCHWARTZ S H,TAN B C,GAGE D A,et al.Specific oxidative cleavage of carotenoids by VP14 of maize[J].Science,1997,276:1872-1874. [25]FRUSCIANTE S,DIRETTO G,BRUNO M,et al.Novel carotenoid cleavage dioxygenase catalyzes the first dedicated step in saffron crocin biosynthesis[J].Proceedings of the National Academy of Sciences,2014,111:12246-12251. [26]LI L,YUAN H.Chromoplast biogenesis and carotenoid accumulation[J].Arch Biochem Biophys,2013,539:102-109. [27]FRASER P D,TRUESDALE M R,BIRD C R,et al.Carotenoid biosynthesis during tomato fruit development(evidence for tissue-specific gene expression)[J].Plant Physiology,1994,105:405-413. [28]HUGUENEY P,BOUVIER F,BADILLO A,et al.Developmental and stress regulation of gene expression for plastid and cytosolic isoprenoid pathways in pepper fruits[J].Plant Physiology,1996,111:619-626. [29]LIU Q,XU J,LIU YZ,et al.A novel bud mutation that confers abnormal patterns of lycopene accumulation in sweet orange fruit(Citrus sinensis L.Osbeck)[J].Journal of Experimental Botany,2007,58:4161-4171. [30]LYU P,LI N,LIU H,et al.Changes in carotenoid profiles and in the expression pattern of the genes in carotenoid metabolisms during fruit development and ripening in four watermelon cultivars[J].Food Chemistry,2015,174:52-59. [31]TOLEDO-ORTIZ G,HUQ E,RODRIGUEZ-CONCEPCION M.Direct regulation of phytoene synthase gene expression and carotenoid biosynthesis by phytochrome-interacting factors[J].Proceedings of the National Academy of Sciences,2010,107:11626-11631. [32]WELSCH R,MAASS D,VOEGEL T,et al.Transcription factor RAP2 and its interacting partner SINAT2:stable elements in the carotenogenesis of Arabidopsis leaves[J].Plant Physiology,2007,145(3):1073-1085. [33]VREBALOV J,RUEZINSKY D,PADMANABHAN V,et al.A MADS-box gene necessary for fruit ripening at the tomato ripening-inhibitor(rin)locus[J].Science,2002,296:343-346. [34]FUJISAWA M,NAKANO T,SHIMA Y,et al.A large-scale identification of direct targets of the tomato MADS box transcription factor RIPENING INHIBITOR reveals the regulation of fruit ripening[J].Plant Cell,2013,25:371-386. [35]LUO Z D,ZHANG J H,LI J,et al.A STAY-GREEN protein SlSGR1 regulates lycopene and β-carotene accumulation by interacting directly with SlPSY1 during ripening processes in tomato[J].New Phytologist,2013,198:442-452. [36]JANELLE M S,LAUREN E S,AMY M L,et al.An R2R3-MYB transcription factor regulates carotenoid pigmentation in Mimulus lewisii flowers[J].New Phytologist,2015,209:1049-1057. [37]CAMPISI L,FAMBRINI M,MICHELOTTI V,et al.Phytoene accumulation in sunflower decreases the transcript levels of the phytoene synthase gene[J].Plant Growth Regulation,2006,48:79-87. [38]KACHANOVSKY D E,FILLER S,ISAACSON T,et al.Epistasis in tomato color mutations involves regulation of phytoene synthase 1 expression by cis-carotenoids[J].Proceedings of the National Academy of Sciences,2012,46:19021-19026. [39]AL-BABILI S,VONLINTIG J,HAUBRUCK H,et al.A novel,soluble form of phytoene desaturase from Narcissus pseudonarcissus chromoplasts is Hsp70-omplexed and competent for flavinylation,membrane association and enzymatic activation[J].Plant Journal,1996(9):601-612. [40]WELSCH R,BEYER P,HUGUENEY P,et al.Regulation and activation of phytoene synthase,a key enzyme in carotenoid biosynthesis,during photomorphogenesis[J].Planta,2000,211:846-854. [41]ZHOU X J,WELSCH R,YANG Y,et al.Arabidopsis OR proteins are the major posttranscriptional regulators of phytoene synthase in controlling carotenoid biosynthesis[J].Proceedings of the National Academy of Sciences,2015,112:3558-3563. [42]LIU Y,ROOF S,YE Z,et al.Manipulation of light signal transduction as a means of modifying fruit nutritional quality in tomato[J].Proceedings of the National Academy of Sciences,2004,101:9897-9902. [43]GALPAZ N,WANG Q,MENDA N,et al.Abscisic acid deficiency in the tomato mutant high-pigment 3 leading to increased plastid number and higher fruit lycopene content[J].Plant Journal,2008,53:717-730. [44]LU S,van ECK J,ZHOU X,et al.The cauliflower Or gene encodes a DnaJ cysteine-rich domain-containing protein that mediates high levels of β-carotene accumulation[J].The Plant Cell,2006,18(12):3594-3605. [45]LOPEZ A B,van ECK J,CONLIN B J,et al.Effect of the cauliflower Or transgene on carotenoid accumulation and chromoplast formation in transgenic potato tubers[J].Journal of Experimental Botany,2008,59:213-223. [46]LI L,YANG Y,XU Q,et al.The Or gene enhances carotenoid accumulation and stability during post-harvest storage of potato tubers[J].Molecular Plant,2012(5):339-352. [47]KIM S H,AHN Y O,AHN M J,et al.Cloning and characterization of an Orange gene that increases carotenoid accumulation and salt stress tolerance in transgenic sweetpotato cultures[J].Plant Physiology and Biochemistry,2013,70:445-454. [48]NETA-SHARIR I,ISAACSON T,LURIE S,et al.Dual role for tomato heat shock protein 21:protecting photosystem II from oxidative stress and promoting color changes during fruit maturation[J].The Plant Cell,2005(17):1829-1838. [49]CHIOU C Y,PAN H A,CHUANG Y N,et al.Differential expression of carotenoid-related genes determines diversified carotenoid coloration in floral tissues of Oncidium cultivars[J].Planta,2010,232(4):937-948. [50]HAN Y,WANG X,CHEN W,et al.Differential expression of carotenoid-related genes determines diversified carotenoid coloration in flower petal of Osmanthus fragrans[J].Tree Genetics & Genomes,2013,10(2):329-338. [51]CAZZONELLI C I,CUTTRISS A J,COSSETTO S B,et al.Regulation of carotenoid composition and shoot branching in Arabidopsis by a chromatin modifying histone methyltransferase,SDG8[J].Plant Cell,2009,21(1):39-53. [52]FU X M,FENG C,WANG C Y,et al.Involvement of multiple phytoene synthase genes in tissue and cultivar-specific accumulation of carotenoids in loquat[J].Journal of Experimental Botany,2014,63:4679-4689. [53]ZHANG B,LIU C,WANG Y Q,et al.Disruption of a CAROTENOID CLEAVAGE DIOXYGENASE 4 gene converts flower colour from white to yellow in Brassica species[J].New Phytologist,2015(4):1513-1526. [54]TZURI G,ZHOU X,CHAYUT N,et al.A‘golden’SNP in CmOr governs fruit flesh color of melon(Cucumis melo)[J].Plant Journal,2015,82:267-279. [55]WELSCH R,ARANGO J,BAR C,et al.Provitamin A accumulation in cassava(Manihot esculenta)roots driven by a single nucleotide polymorphism in a phytoene synthase gene[J].Plant Cell,2010(22):3348-3356. [56]HA S H,KIM J B,PARK J S,et al.A comparison of the carotenoid accumulation in Capsicum varieties that show different ripening colours:deletion of the capsanthin-capsorubin synthase gene is not a prerequisite for the formation of a yellow pepper[J].Journal of Experimental Botany,2007,58(12):3135-3144. [57]NAZIA N,LI L,SHAN L,et al.Carotenoid Metabolism in Plants[J].Molecular Plant,2015(8):68-82.