[1]KNUDSEN J T,TOLLSTEN L,BERGSTRM L G.Floral scents-a checklist of volatile compounds isolated by head-space techniques[J].Phytochemistry,1993,33(2):253-280. [2]THOLL D,KISH C M,ORLOVA I,et al.Formation of monoterpenes in Antirrhinum majus and Clarkia breweri flowers involves heterodimeric geranyl diphosphate synthases[J].The Plant Cell,2004,16(4):977-992. [3]BOATRIGHT J,NEGRE F,CHEN X,et al.Understanding in vivo benzenoid metabolism in petunia petal tissue[J].Plant Physiology,2004,135(4):1993-2011. [4]DUDAREVA N,PICHERSKY E.Metabolie engineering of floral scent of ornamentals[J].Journal of Crop Improvement,2006,18(1-2):325-346. [5]CHANDLER S F,SANCHEZ C.Genetic modification,the development of transgenic ornamental plant varieties[J].Plant Biotechnology Journal,2012,10(8):891-903. [6]GERATS T,VANDENBUSSCHE M.A model system for comparative research:Petunia[J].Trends in Plant Science,2005,10(5):251-256. [7]SCHUURINK R C,HARING M A,CLARK D G.Regulation of volatile benzenoid biosynthesis in petunia flowers[J].Trends in Plant Science,2006,11(1):20-25. [8]TANAKA Y,SASAKI N,OHMIYA A.Biosynthesis of plant pigments:anthocyanins,betalains and carotenoids[J].The Plant Journal,2008,54(4):733-749. [9]DUDAREVA N,PICHERSKY E.Biology of floral scent[M].Boca Raton:CRC Press,2006. [10]JONES R L,BUCHANAN B B,GRUISSEM W.Biochemistry and molecular biology of plants[M].Wiley Press,2002. [11]DUDAREVA N,CSEKE L,BLANC V M,et al.Evolution of floral scent in Clarkia:novel patterns of S-linalool synthase gene expression in the C.breweri flower[J].The Plant Cell,1996,8(7):1137-1148. [12]PICHERSKY E,GANG D R.Genetics and biochemistry of secondary metabolites in plants:an evolutionary perspective[J].Trends in Plant Science,2000,5(10):439-445. [13]PICHERSKY E,NOEL J P,DUDAREVA N.Biosynthesis of plant volatiles:nature’s diversity and ingenuity[J].Science,2006,311(5762):808-811. [14]SCHWAB W.Metabolome diversity:too few genes,too many metabolites?[J].Phytochemistry,2003,62(6):837-849. [15]WU S,CHAPPELL J.Metabolic engineering of natural products in plants;tools of the trade and challenges for the future[J].Current Opinion in Biotechnology,2008,19(2):145-152. [16]THOLL D.Terpene synthases and the regulation,diversity and biological roles of terpene metabolism[J].Current Opinion in Plant Biology,2006,9(3):297-304. [17]FERRER J,AUSTIN M B,STEWART C,et al.Structure and function of enzymes involved in the biosynthesis of phenylpropanoids[J].Plant Physiology and Biochemistry,2008,46(3):356-370. [18]TZIN V,GALILI G.New insights into the shikimate and aromatic amino acids biosynthesis pathways in plants[J].Molecular Plant,2010,3(6):956-972. [19]VERDONK J C,de VOS C R,VERHOEVEN H A,et al.Regulation of floral scent production in petunia revealed by targeted metabolomics[J].Phytochemistry,2003,62(6):997-1008. [20]DIXON R A,CHEN F,GUO D,et al.The biosynthesis of monolignols:a “metabolic grid”,or independent pathways to guaiacyl and syringyl units?[J].Phytochemistry,2001,57(7):1069-1084. [21]COLQUHOUN T A,KIM J Y,WEDDE A E,et al.PhMYB4 fine-tunes the floral volatile signature of Petunia×hybrida through PhC4H[J].Journal of Experimental Botany,2010:q342. [22]UNDERWOOD B A,TIEMAN D M,SHIBUYA K,et al.Ethylene-regulated floral volatile synthesis in petunia corollas[J].Plant Physiology,2005,138(1):255-266. [23]VERDONK J C,HARING M A,van TUNEN A J,et al.ODORANT1 regulates fragrance biosynthesis in petunia flowers[J].The Plant Cell,2005,17(5):1612-1624. [24]NEGRE F,KISH C M,BOATRIGHT J,et al.Regulation of methylbenzoate emission after pollination in snapdragon and petunia flowers[J].The Plant Cell,2003,15(12):2992-3006. [25]MAEDA H,SHASANY A K,SCHNEPP J,et al.RNAi suppression of Arogenate Dehydratase1 reveals that phenylalanine is synthesized predominantly via the arogenate pathway in petunia petals[J].The Plant Cell,2010,22(3):832-849. [26]COLQUHOUN T A,VERDONK J C,SCHIMMEL B C,et al.Petunia floral volatile benzenoid/phenylpropanoid genes are regulated in a similar manner[J].Phytochemistry,2010,71(2):158-167. [27]COLQUHOUN T A,SCHIMMEL B C,KIM J Y,et al.A petunia chorismate mutase specialized for the production of floral volatiles[J].The Plant Journal,2010,61(1):145-155. [28]KOLOSOVA N,SHERMAN D,KARLSON D,et al.Cellular and subcellular localization of S-Adenosyl-l-Methionine:Benzoic acid carboxyl methyltransferase,the enzyme responsible for biosynthesis of the volatile ester methylbenzoate in snapdragon flowers[J].Plant Physiology,2001,126(3):956-964. [29]BOATRIGHT J,NEGRE F,CHEN X,et al.Understanding in vivo benzenoid metabolism in petunia petal tissue[J].Plant Physiology,2004,135(4):1993-2011. [30]KOEDUKA T,FRIDMAN E,GANG D R,et al.Eugenol and isoeugenol,characteristic aromatic constituents of spices,are biosynthesized via reduction of a coniferyl alcohol ester[J].Proceedings of the National Academy of Sciences,2006,103(26):10128-10133. [31]DEXTER R,QUALLEY A,KISH C M,et al.Characterization of a petunia acetyltransferase involved in the biosynthesis of the floral volatile isoeugenol[J].The Plant Journal,2007,49(2):265-275. [32]COLQUHOUN T A,MARCINIAK D M,WEDDE A E,et al.A peroxisomally localized acyl-activating enzyme is required for volatile benzenoid formation in a Petunia×hybrida cv.‘Mitchell Diploid’flower[J].Journal of Experimental Botany,2012,63(13):48214833. [33]ZVI M M B,NEGRE Z F,MASCI T,et al.Interlinking showy traits:co-engineering of scent and colour biosynthesis in flowers[J].Plant Biotechnology Journal,2008,6(4):403-415. [34]SPITZER-RIMON B,MARHEVKA E,BARKAI O,et al.EOBII,a gene encoding a flower-specific regulator of phenylpropanoid volatiles’ biosynthesis in petunia[J].The Plant Cell,2010,22(6):1961-1976. [35]COLQUHOUN T A,CLARK D G.Unraveling the regulation of floral fragrance biosynthesis[J].Plant Signaling and Behavior,2011,6(3):378-381. [36]SPITZER-RIMON B,FARHI M,ALBO B,et al.The R2R3-MYB-like regulatory factor EOBI,acting downstream of EOBII,regulates scent production by activating ODO1 and structural scent-related genes in petunia[J].The Plant Cell,2012,24(12):5089-5105. [37]van MOERKERCKE A,GALVAN-AMPUDIA C S,VERDONK J C,et al.Regulators of floral fragrance production and their target genes in petunia are not exclusively active in the epidermal cells of petals[J].Journal of Experimental Botany,2012,63(8):3157-3171.